Chapter 16

Calculus for the Life Sciences: A Modeling Approach Volume II · 11 exercises

Problem 4

Examine the behavior of \(x_{t}\) for \(t=2,3, \cdots, 20\) of the difference equation $$ \begin{aligned} x_{t+1} &=\alpha_{0} x_{t}+\alpha_{1} x_{t-1} \\ x_{0} &=0.1, \quad x_{1}=0.2 \end{aligned} $$ for a. \(\alpha_{0}=\frac{5}{6}, \quad \alpha_{1}=-\frac{1}{2}\). b. \(\alpha_{0}=\frac{3}{2}, \quad \alpha_{1}=-\frac{1}{2}\). c. \(\alpha_{0}=\frac{5}{2}, \quad \alpha_{1}=-1\).

7 step solution

Problem 5

For the symbiosis system, $$ \begin{aligned} x_{n+1}-x_{n} &=\frac{5}{98} \cdot x_{n}\left(1+0.4 y_{n}-x_{n}\right) \\ y_{n+1}-y_{n} &=\frac{7}{120} \cdot y_{n}\left(1+\frac{5}{7} x_{n}-y_{n}\right) \end{aligned} $$ (1.76,2.0) is an equilibrium point. Choose the initial value, \(\left(x_{0}, y_{0}\right)=(1.76,2.1)\) and compute \(\left(x_{1}, y_{1}\right),\) \(\left(x_{2}, y_{2}\right),\left(x_{3}, y_{3}\right) .\)

6 step solution

Problem 5

For the dynamical system \(16.6 \mathrm{C}\), $$ \begin{array}{lll} x_{0} & =0.25 & x_{n+1} & =0.9 \times x_{n}+0.04 \times y_{n} \\ y_{0} & =0.5 & y_{n+1} & =0.1 \times x_{n}+0.86 \times y_{n} \end{array} $$ the characteristic roots are \(r_{1}=0.946\) and \(r_{2}=0.814 . x_{t}\) and \(y_{t}\) are given by $$ x_{t}=C_{1} \times r_{1}^{t}+C_{2} \times r_{2}^{t} $$ $$ y_{t}=D_{1} \times r_{1}^{t}+D_{2} \times r_{2}^{t} $$ Where \(C_{1}, C_{2}, D_{1}\) and \(D_{2}\) are computed from $$ \begin{array}{ll} x_{0}=C_{1}+C_{2} & y_{0}=D_{1}+D_{2} \\ x_{1}=C_{1} \times r_{1}+C_{2} \times r_{2} & y_{1}=D_{1} \times r_{1}+D_{2} \times r_{2} \end{array} $$ Compute \(C_{1}, C_{2}, D_{1}\) and \(D_{2}\) and use these values in Equations 16.12 and 16.13 to compute \(x_{2}, x_{3},\) and \(y_{2}, y_{3}\)

4 step solution

Problem 6

For the dynamical system \(16.6 \mathrm{D}\), $$ \begin{array}{lll} x_{0} & =0.25 & x_{n+1}= & 1.15 \times x_{n}-0.8 \times y_{n} \\ y_{0} & =0.5 & y_{n+1}= & 0.1 \times x_{n}+0.86 \times y_{n} \end{array} $$ the characteristic roots are $$ r_{1} \doteq 1.005+0.243 i, \quad r_{2} \doteq 1.005-0.243 i $$ \(x_{t}\) and \(y_{t}\) are given by $$ \begin{aligned} x_{t} &=C_{1} \rho^{t} \cos t \theta+C_{2} \rho^{t} \sin t \theta \\ y_{t} &=D_{1} \rho^{t} \cos t \theta+D_{2} \rho^{t} \sin t \theta \end{aligned} $$ where \(\rho=\sqrt{1.005^{2}+0.243^{2}}, \theta=\arccos (1.005 / \rho)\) and \(C_{1}, C_{2}, D_{1}\) and \(D_{2}\) are computed from $$ \begin{array}{ll} x_{0}=C_{1} & y_{0}=D_{1} \\ x_{1}=C_{1} \rho \cos \theta+C_{2} \rho \sin \theta & y_{1}=D_{1} \rho \cos \theta+D_{2} \rho \sin \theta \end{array} $$ Compute \(\rho, \theta\) and \(C_{1}, C_{2}\) and \(D_{1}, D_{2}\) and use these values in Equations 16.14 and 16.14 to compute \(x_{2},\) \(x_{3},\) and \(y_{2}, y_{3}\)

7 step solution

Problem 7

For the following systems, find all of the equilibrium points \(\left(x_{e}, y_{e}\right)\). a. For each equilibrium point with \(x_{e}>0\) and \(y_{e}>0,\) determine the stability of the system at that equilibrium point. b. Draw a phase plane and in each region of the phase plane bounded by null lines, draw a vector pointing from a point \(\left(x_{n}, y_{n}\right)\) toward \(\left(x_{n+1}, y_{n+1}\right)\). \(\begin{aligned} \text { a. } & x_{n+1}-x_{n}=0.1 * x_{n} *\left(1-0.5 y_{n}-x_{n}\right) \\ & y_{n+1}-y_{n}=0.05 * y_{n} *\left(1+0.2 x_{n}-y_{n}\right) \\ \text { b. } & x_{n+1}-x_{n}=0.05 * x_{n} *\left(1-0.2 y_{n}-x_{n}\right) \\ & y_{n+1}-y_{n}=0.1 * y_{n} *\left(1-0.5 x_{n}-y_{n}\right) \\ \text { c. } \quad & x_{n+1}-x_{n}=0.2 * x_{n} *\left(1+0.4 y_{n}-x_{n}\right) \\ & y_{n+1}-y_{n}=0.4 * y_{n} *\left(1+0.8 x_{n}-y_{n}\right) \\ \text { d. } \quad & x_{n+1}-x_{n}=0.05 * x_{n} *\left(1-0.2 y_{n}-x_{n}\right) \\ & y_{n+1}-y_{n}=0.1 * y_{n} *\left(1-y_{n}\right) \end{aligned}\)

5 step solution

Problem 7

Determine whether the following systems are stable. a. \(\quad \begin{aligned} x_{t+1} &=0.48 x_{t}+0.2 y_{t} \\ y_{t+1} &=0.128 x_{t}+0.72 y_{t} \end{aligned}\) b. \(\quad \begin{aligned} x_{t+1} &=0.8 x_{t}-0.5 y_{t} \\ y_{t+1} &=0.2 x_{t}+0.9 y_{t} \end{aligned}\) c. \(\quad \begin{aligned} x_{t+1} &=1.1 x_{t}-0.5 y_{t} \\ y_{t+1} &=0.2 x_{t}+0.9 y_{t} \end{aligned}\) d. \(\quad \begin{aligned} x_{t+1} &=0.8 x_{t}-0.1 y_{t} \\ y_{t+1} &=0.1 x_{t}+0.6 y_{t} \end{aligned}\) e. \(\quad \begin{aligned} x_{t+1} &=1.2 x_{t}-0.1 y_{t} \\ y_{t+1} &=0.9 x_{t}+0.6 y_{t} \end{aligned}\) f. \(\quad \begin{aligned} x_{t+1} &=0.56 x_{t}+0.4 y_{t} \\ y_{t+1} &=0.256 x_{t}+1.04 y_{t} \end{aligned}\)

9 step solution

Problem 11

In Equation 16.11 , we claim that for $$ x_{t}=C_{1} r_{1}^{t}+C_{2} \times t \times r_{1}^{t}, \quad \lim _{t \rightarrow \infty} A_{t}=0 \quad \text { if } \quad\left|r_{1}\right|<1 $$ Use L'Hospital's Theorem 14.5.1 to show that \(\lim _{t \rightarrow \infty} t \times r^{t}=0\) if \(|r|<1\).

5 step solution

Problem 12

Show that the characteristic equation of $$ \begin{aligned} x_{n+1} &=p \cdot x_{n}-q \cdot y_{n} \\ y_{n+1} &=x_{n} \end{aligned} $$ is Exercise 16.2 .12 Show that the characteristic equation of $$ \begin{array}{l} x_{n+1}=p \cdot x_{n}-q \cdot y_{n} \\ y_{n+1}=x_{n} \end{array} $$ is $$ x_{n+2}-p x_{n+1}+q x_{n}=0 $$

6 step solution

Problem 17

Compute the characteristic values of $$ \begin{array}{l} \xi_{n+1}=0.7647 \xi_{n}+-0.04706 \eta_{n} \\ \eta_{n+1}=0.02157 \xi_{n}+0.7843 \eta_{n} \end{array} $$ and determine whether both of them are of magnitude less than \(1 .\)

6 step solution

Problem 18

For each of the following systems, find an equilibrium point which has both coordinates positive, if there is one, and determine whether it is stable. $$ \begin{array}{l} \text { a. } \quad x_{n+1}=x_{n}+0.7 x_{n}\left(1-0.5 y_{n}-x_{n}\right) \quad y_{n+1}=y_{n}+0.3 y_{n}\left(1+0.2 x_{n}-y_{n}\right) \\ \text { b. } x_{n+1}=x_{n}+0.7 x_{n}\left(1-0.8 y_{n}-x_{n}\right) \quad y_{n+1}=y_{n}+0.3 y_{n}\left(1+0.5 x_{n}-y_{n}\right) \end{array} $$ c. \(x_{n+1}=x_{n}+1.2 x_{n}\left(1-0.9 y_{n}-x_{n}\right) \quad y_{n+1}=y_{n}+0.8 y_{n}\left(1+0.2 x_{n}-y_{n}\right)\) d. \(x_{n+1}=x_{n}+1.8 x_{n}\left(1-1.2 y_{n}-x_{n}\right) \quad y_{n+1}=y_{n}+1.2 y_{n}\left(1+0.5 x_{n}-y_{n}\right)\) e. \(\quad x_{n+1}=x_{n}+1.4 x_{n}\left(1-1.0 y_{n}-x_{n}\right) \quad y_{n+1}=y_{n}+0.8 y_{n}\left(1+0.5 x_{n}-y_{n}\right)\) f. \(x_{n+1}=x_{n}+1.4 x_{n}\left(1-1.1 y_{n}-x_{n}\right) \quad y_{n+1}=y_{n}+0.8 y_{n}\left(1+0.5 x_{n}-y_{n}\right)\)

7 step solution

Problem 21

For each of the following systems, find an equilibrium point which has both coordinates positive, if there is one, and determine whether it is stable. $$ \begin{array}{lll} \text { a. } & x_{n+1}=x_{n}+0.7 x_{n}\left(1-0.5 y_{n}-x_{n}\right) & y_{n+1}=y_{n}+0.3 y_{n}\left(1-0.2 x_{n}-y_{n}\right) \\ \text { b. } & x_{n+1}=x_{n}+0.7 x_{n}\left(1-0.8 y_{n}-x_{n}\right) & y_{n+1}=y_{n}+0.3 y_{n}\left(1-0.5 x_{n}-y_{n}\right) \\ \text { c. } & x_{n+1}=x_{n}+1.2 x_{n}\left(1-0.9 y_{n}-x_{n}\right) & y_{n+1}=y_{n}+0.8 y_{n}\left(1-0.2 x_{n}-y_{n}\right) \\ \text { d. } & x_{n+1}=x_{n}+1.8 x_{n}\left(1-1.2 y_{n}-x_{n}\right) & y_{n+1}=y_{n}+1.2 y_{n}\left(1-0.5 x_{n}-y_{n}\right) \\ \text { e. } & x_{n+1}=x_{n}+1.4 x_{n}\left(1-1.0 y_{n}-x_{n}\right) & y_{n+1}=y_{n}+0.8 y_{n}\left(1-0.5 x_{n}-y_{n}\right) \\ \text { f. } & x_{n+1}=x_{n}+1.4 x_{n}\left(1-0.9 y_{n}-x_{n}\right) & y_{n+1}=y_{n}+0.8 y_{n}\left(1-0.5 x_{n}-y_{n}\right) \end{array} $$

8 step solution

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